,      Brachytheciaceae, Dicranaceae  Mniaceae,        .   ,   « »  195  (165    30 ). ,     33%   . 
 ..,  ..      . Sphagnaceae ­ Hedwigiaceae. ., 2003. . 1. 608 .  ..,  ..      . Fontinaceae ­ Amblystegiaceae. ., 2004. . . 2. 609­944.    ,      ( .  ..  .). : . . . -, 1993. 210 .    . .: , 1980. 429 .     « » / ..   .;  . .. . .: . .       , 2004. 62 . [    ; . 4].

               . .*,  . .**,  . .***
 - . ..  , . -, . * Paul@PK15777.spb.edu, **vvgor@skylink.spb.ru, ***nata@AS7542.spb.edu

       ,  ,        (, , 1983).                 (, , , 2005).               ,      .          (   45  55 )    (  220, 260, 320   400 ).          
236

 ,        .      (      1.3   4 )      15%  ,   ­ 60­90%.   ,         (   1.3     1.3   0.1  4 )   (   1.3 )     (  30%  ).         .        .         ,    .         4           .        ­1 (, , , 2005).             5   .          45 ,   1-   ,      ­  (   1.3     22 ) ­         .        5     0.040  0.200 /     0.098 /.    37%.  ,       55 ,      ,    .    (  19 )   (  41 )   0.056  0.256     0.120  0.156 /    5  .    (  50 )   0.062  0.488 /   5       0.242 .           56%.      220     (  19 )  5     0.050  0.220 / (  0.114),  (  125 ) ­  0.040  0.560 / (  0.244),   (  187 ) ­  0.102  0.940 / (  0.380).         48%.    260      (   41 ,  ­ 79
237

)    0.030­0.300 / (    0.094  0.138 ).    (  174 )   0.040  0.840     0.232 /    5  .       63%.       320 ,      ,             (  37 )   (  102 ).      0.030  0.200      0.095 /.     ( 196 )  0.194      0.048­0.640 /  5 .        68%.        400         (  20 )   0.040  0.360 /     0.128 /   5  .      (  73 )     0.040  0.844      0.196 /.    (  152 )       0.066­0.700 / (  0.234 ).         72%.               5         : «» < «» < « ».   ,    ,     ~1.5 ,    ,   ~2.5 ,   .                  .         .                 :  ,    ,    .         .       ,               (, , 1973; , 1983).
238

   .    ,        ,             .           . ,      45­220  260­400           (  t=3.39    <0.05)   41  51% .         (t=2.73, <0.05)      45­55  37­45% (  42%),    220­400  ­ 40­74% (  55%).  ,         ,       ,     .         ,          .          ,   ,                       (Steijlen, Zackrisson, 1987; Drobyshev, 2001).               ,       .      : 1)                     0.030  0.940 / (   0.094  0.380 /). 2)               2   37% (      45 )  72% (       400 ). 3)           5      ­  :  :   ­    ­ 2.5 : 1.5 : 1 ­       .
239


 ..,  ..       //     . .: , 1983. . 65­78.  ..,  ..        //       . .: , 1973. . 242­246.  ..,  ..,  ..      ,       Picea obovata Ledeb.  Betula pubescens Ehrh.            //     . ., 2005. 256­272 .  ..           // . 1983.  3. . 71­79. Drobyshev I.V. Effect of natural disturbances on the abundance of Norway spruce (Picea abies (L.) Karst.) regeneration in nemoral forests of the southern boreal zone // For. Ecol. Manage. 2001. Vol. 140. P. 151­161. Engelmark O. Early post-fire tree regeneration in a Picea-Vaccinium forest in northern Sweden // J. Veg. Sci. 1993. Vol. 4. P. 791­794. Steijlen I., Zackrisson O. Long-term regeneration dynamics and successional trends in northern Swedish coniferous forest stand // Can. J. Bot. 1987. Vol. 65. P. 839­848.

              . .
    .. , . , . Yulia 1975@list.ru

              .    25         .         .          ,        (  ., 2001).                ­     . ,      ,    240