, , . . . ­ 72 , (, , ) . . . . . . , . . ­ . . . . .. , . , . kovalyovy@mail.ru () . 252 (=280­320 ) (Huttunen, Laakso, 1998). - (Teramura, Sullivan, 1994; Fiscus, Booker, 1995; Fritzemeier, Kindl, 1981). , (Teramura, Sullivan, 1994; Fiscus, Booker, 1995). , (Fritzemeier, Kindl, 1981). 100 . ( ) , (Tobin, Silverthorne, 1985; Jordan, James, Strid, Anthony, 1994; Fuglevand, Jackson J.A., Jenkins G.I., 1996). . 14- , -400 -400 ( ­ 16/8 ) 20±2°. - (++) 1 = 120 /2 (10 .); 2 = 240 /2 (20 .); = 360 /2 (30 .). - -81. - -1000. , . , , .., .. 1986 . -26. 70%- . 253 .., .. 3­ 5- . ( ) , , / , -, . , ­ ­ (Stafford, 1991). , , ( ) , (Jordan, James, Strid, Anthony, 1994). , 120 /2 (% ) 109% ; 240 360 /2 125% 59% . , , , , (, ). , (3­4 24­48 ) . (Teramura, Sullivan, 1994; Fiscus, Booker, 1995) , de novo (, , ) , 254 , ( ) , -, (, , , 2002) . /- - ­ , - (, 1969), (Sutherland, 1981) - (, UVB-) (Fuglevand, Jackson, Jenkins, 1996). , ( , , .), (), (, , , 2002) (Sävenstrand, Broshé, Strid, 2002; , , 2002) (/-) . .., .., .. // . . . . 2002. . 34. 3. . 187­198. .. . .: , 1968. 250 . .. // . . 1987. . 34. . 4. . 698­711. .., .. // . . 1986. . 33. 5. . 1013­1026. .. . .: , 1969. 511 . .., .. // . . . . 2002. . 34. 3. . 213­226. .., .. // . 1991. . 29. . 173­177. 255 Fiscus E.L., Booker F.L. Is increased UV-B a threat to crop photosynthesis and productivity // Photosynth. Res. 1995. Vol. 43. P. 81­92. Fritzemeier K.H., Kindl H. Coordinate induction by UV light of stilbene synthase, phenylalanine ammonialyase and cinnamate 4-hydroxylase in leaves of Vitaceae // Planta. 1981. Vol. 151, N 1. P. 48­52. Fuglevand G., Jackson J.A., Jenkins G.I. UV-B, UV-A and blue light signal transduction pathways interact synergistically to regulate chalcone synthase gene expession in Arabidopsis // Plant Cell. 1996. Vol. 8. P. 2347­2357. Huttunen S.K.H., Laakso K. Impact of increased UV-B on plant ecosystems // Chemosphere. 1998. Vol. 36. P. 829­833. Jordan B.R., James P.E., Strid A., Anthony R.G. The effect of ultraviolet-B radiation on gene expression and pigment composition in etiolated and green pea leaf tissue: UV-B induced changes are gene-specific and dependent upon the development stage. // Plant Cell Environ. 1994. Vol. 17. P. 45­54. Sävenstrand H., Broshé M., Strid A. Regulation of gene expression by low levels of ultraviolet-B radiation in Pisum sativum: Isolation of novel genes by suppression subtractive hybridization // Plant and Cell Physiol. 2002. Vol. 43, N 4. P. 402­410. Stafford H.A. Flavonoid evolution: an enzymic approach // Plant Physiol. 1991. Vol. 96. P. 680­685. Sutherland B.M. Photoreactivation // BioScience. 1981. Vol. 31. P. 439­444. Teramura A.H., Sullivan J.H. Effects of UV-B radiation on photosynthesis and growth of terrestrial plants // Photosynth. Res. 1994. Vol. 39. P. 463­473. Tobin E.M., Silverthorne J. Light regulation of gene expression in higher plants // Annu. Rev. Plant Physiol. 1985. Vol. 36. P. 569­593. . .*, . . **, . .*, . .*, . .*, . .*, . .*, . .*, . .* . .. , . -, . oxyria@gmail.com. ** , . , . zverev@ecos.tsu.ru * . .. 50 () ( ., 2001), 148 . IBIS (, 256